Wednesday, September 17, 2014

Primates

        The lemur is a primitive primate that can be found in Madagascar. The term Lemur not only applies to the typical lemurs but also to the avahis, sifakas, indri, and aye-aye of Madagascar. The island country of Madagascar is located on the southeastern coast of Africa. The island consist of three parallel longitudinal zones-the central plateau, the coastal strip on the east side, and the low plateaus and plains in the west. The steep eastern plateau has numerous short, torrential rivers which discharge either into the coastal lagoons or directly into the sea over waterfalls or rapids. The western side of the plateau is crossed by longer and larger rivers, which bring large deposits of fertile alluvium down into the plains. The island at one point was covered with evergreen and deciduous forest, but now only little remains except on the eastern escarpment and in some areas in the west. The forest was cut in order to clear for rice fields, to obtain building materials, and to export valuables such as ebony, rosewood, and sandalwood. A small part of the island is covered with prairie grasses and bamboo or small thin trees. The south part of the island is arid and grows thorn trees, giant cacti, dwarf baobab trees, succulents and other drought-resistant plants. Due to the island's isolation many primitive primates have survived and evolved into unique forms. It's been estimated that about 40 species of lemurs live on Madagascar.
The lemur has a unique trait called a "dental comb", which is formed by forward projecting lower incisors and canines. The modified teeth are used by the lemurs for grooming similar to a hair comb and feeding. The structure of the jaw has evolved independently in different types of mammals through convergent evolution. In most mammals the toothcombs include incisors only, but in lemuriform primates the jaw includes incisors and canine teeth that tilt forward by a canine-shaped first premolar. The toothcomb can also be used for food procurement and bark gouging. Due to the islands many plants, trees and shrubs, the lemurs jaw has adapted to the surrounding environment. The lemur has a wide variety of foods such as leaves, fruits, buds, bark and shoots. Lemurs are also known to eat insects and even small animals when fruits are not in season, and when conditions are really harsh lemur's hibernate which helps contributes to their survival. The lemur has a dental formula of 2.1.3.3, and has  reduced incisors and tooth comb when compared to the spider monkey.


       The spider monkey is a smaller primate that can be found in the forest areas of southern Mexico through central and South America to brazil. The habitat of the spider monkey consist of tropical rainforests, evergreen, semi deciduous and mangrove forest. The current forest where the spider monkey's are located is in danger due to the removal of the forest to make way for new farm land. The spider monkeys are well adapted to the high treetops as they are strong and agile. The spider monkey has a wide variety of foods, such as nuts, fruits, leaves, bird eggs, spiders and other insects. The spider monkey or new world monkey, has a dental formula of 2.1.3.3 (two incisors, one canine, three premolars, and three molars). The maxilla and mandible of the monkey are equally the same, and the assortment of teeth allows the primates to have a diverse diet. The incisors teeth of the monkey help with cutting into food, and the canines are stabbing teeth that can be used as a weapon. The premolars and molars are mainly used for shearing and crushing food. The spider monkey has a unique dental formula which has evolved due to the environment the primate lives in. The diet of the spider monkey has also contributed to the evolution of the primates teeth, which is due to the lack of dietary specialization and reflects the shape and size of the primates teeth. This type of dentition was created from the varied diet of the primate, and has contributed to their overall success during the last 50 million years.


        The next primate is the baboon also known as the Old World Monkey. There are 5 different species of baboons and each hierarchical troop of baboons can have up to 300 members. Baboons can be found in the savanna and semiarid environments in Africa to Arabia. Very few baboons can be found in tropical forests. They have also been spotted in some open country parts of Africa in the Sahara southward, and in southern Africa south of the Zambezi River. These locations consist of a desert environment which is common throughout the Sahara. The Sahara is bordered by the Atlantic Ocean, Atlas Mountains, Red Sea and Mediterranean Sea. The baboon eats a variety of plants and animals which includes grass and grass seed, fruits, pods, roots, and tubers. They are also known to eat rodents, birds, gazelle fawns, and in some parts of South Africa they kill lambs and are known as crop raiders. The dentition formula for a baboon is 2.1.2.3, which is acquired from the primate having a slightly different diet compared to New World Monkeys. The dental pattern and formation of the jaw are similar to humans with some key changes. The difference between humans and baboons is the more elongate arch form, prominent maxillary canine crowns, and the second and third molar crowns are greater size compared to the first molar crowns. The baboon males have powerful jaws with longer canine teeth, which can be effective weapons in defense against predators and competition with other males. This type of trait was acquired from the environment the baboon lives in. Due to the primate's diet which consist of eating other animals as compared to the New World Monkeys, they have developed the longer canines which help killing the varied animals in their diet. Since the monkey competes with other primates for the same desirable foods  this leads to the killing of other monkeys, young chimpanzees and even flamingos. Therefore, the dentition formula has evolved into a 2.1.2.3 pattern due the varied diet of the baboon.


      The lesser ape also known as a gibbon, is a smaller ape that is found in the tropical forests of Southeast Asia. The environment where the gibbon can be found is generally composed of broad-leaved trees and wet tropical uplands and lowlands. The rainforests are vegetation type dominated by broad-leaved trees which form dense upper canopies and consist of a diverse vegetation. The gibbons are omnivores, and their diet includes fruit, leaves, flowers, seeds, tree bark, and tender plant shoots. They are also known to eat insects, spiders, bird eggs, and small birds. The gibbon has a skull that closely resembles the great ape. Their teeth also are similar to the great ape, which consist a U-shaped dental arcade and a thin and light mandible. The incisors are broad and flat, while the molars have low, rounded cusps with thick enamel. They also have large, dagger-like canines in both upper and lower jaw. The dentition formula for the gibbons is 2.1.2.3. The trait of the gibbons dentition formula has adapted to its frugivorous environment.


       The Chimpanzee is a species of apes that is most closely related to humans. Chimpanzees can be found in tropical forests and savannas of Africa from The Gambia to Lake Albert, Lake Victoria and Northwestern Tanzania in the east. The tropical forest has a diverse array of vegetation, and broad-leaved trees that form a dense upper canopy. The savanna has vegetation that grows under hot seasonally dry climatic conditions, and has open tree canopies. The chimpanzee consumes many types of fruits, leaves, figs, nuts, flowers, insects and large animals they hunt and kill. Occasionally male and female chimpanzee's take part in group hunting to kill small mammals like bushpigs and antelope. The jaw of a chimpanzee is heavy and protruding, and the canine teeth are large. Male chimpanzees have larger canines compared to females, and use these in battle with other males during predation. The dentition of the chimpanzee is similar to humans except for the larger canines. The dentition formula for chimpanzees is 2.1.2.3/2.1.2.3 which equals 32 teeth. This trait for the chimpanzee dentition formula has been acquired due to their environment and their varied diet.



        Due to the information I have collected it seems that primate dentition is influenced based on the environment they are presented with and their diet. The five groups of primates all have very similar diets, and this could cause competition for food with other species. The overall lack of dietary specialization in primates is reflected and expressed in the size and shape of the teeth, because shape and size are directly related to diet. Most of the primates have premolars and molars with low, rounded cusps which allows them to process most types of food. The primates have developed a dentition that has adapted to a varied diet and has contributed to their overall success for millions of years. 

Sources:
www.Lemurworld.com. BioExpedition Publishing, Web. 15 Sept. 2014
Colin Peter Groves. www.Britannica.com. Web. 16 Sept. 2014
Matthew Tschoegl. learni.st. Web. 15 Sept. 2014
www.angrydmonkey.com. Web. 15 Sept. 2014
Dr. Dennis O'Neil. anthro.palomar.edu. 1998-2014. Web. 16 Sept. 2014

4 comments:

  1. Very detailed post! I'd have to agree with your conclusion that primates teeth have adapted to their environments. I always wonder if our teeth were once more like the chimpanzees or if they were more equipped for eating vegetarian. With all the health information and news emerging in the media lately, I can't help but wonder if our teeth adapted to eating meat originally or if they have adapted to eating meat. Some people protest that meat isn't good for our digestion and I'm curious to know if we adapted to eating meat because other options were scarce or if we were always equipped to eat meat, but modern processing of meat is what we are not supposed to consume.

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  2. Excellent description of the Lemur's environment, and I like how you touched upon the deforestation that can very much so contribute to the extinction of Lemurs. Also I didn't know that baboons could have troops that numbered in the hundreds. I also like that you brought up the comparison between canines of the male and female species of chimpanzees. I agree too that the environment and diets of all the listed primates has influenced their dentition patterns.

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  3. Very good discussion throughout of the environment of each primate. Well done.

    Good connections made between the dental traits of the lemurs and spider monkeys and their environment, primarily of their diet. The shape of the teeth, their cusps, sheering complexes, flat surfaces will all reflect what these primates tend to eat.

    Baboons are a little more difficult because the canine sometimes misleads people, but you did a good job of explaining the non-dietary function of that tooth. But one of the keys to understanding baboon dentition is recognizing just how broad their diet is... it is called "opportunistic", meaning that the eat what they can get and their dentition is generalized to support that important dietary adaptation.

    The loss of the bicuspid tooth in old world monkeys and apes actually has nothing to do with diet. Think about what the jaw is attached to, the upper jaw in particular, and consider how the primate senses and evolved and changed to see if you can figure out why we have not only lost our third bicuspids but also our third molars (that wisdom tooth that can cause so much trouble).

    Good description and connections made for the gibbon dentition.

    Yes, the varied diet of the chimpanzees, much like that of the baboon, have led to the general pattern of dentition to allow for that wide range of food. Very similar to our own general dentition, actually.

    Good post.

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  4. Graycee beat me to it, very detailed post! I really enjoyed reading it because I could visualize each primates environment. I also loved your pictures because it gave me visual proof of each similarity and each difference in all of the primates. I touched briefly on the environment in my post on the locomotor patterns but reading your post gave me a better sense of the way they live and survive which was awesome. Very well written and informative!

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